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Narrator: When they say to watch your back
you really should, because there is
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something pretty important in there called
the Spinal Cord.
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Now, if you think of a nerve as a road
then the spinal cord is huge highway.
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It's a major reflex center and holds many
neural tracks that connect the brain to
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the rest of the body, allowing for
important communication to occur.
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The Spinal Cord starts at the
Foramen Magnum, where it is continuous
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with the Madula Oblongata, which is the
most coddle portion of the brain stem.
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It then extends inferiorly through the
vertebral canal.
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In adults, it usually ends at the level of the first
or second Lumbar vertebra.
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In infants it usually ends at the second
or third Lumbar vertebra.
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The tapered end of the Spinal Cord is
called the Conus Medullaris.
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If we look at a transverse section of the
spinal cord we can see the anterior
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median fissure that extends along the
midline of the Spinal Cord anteriorly,
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similarly, the posterior median sulcus
extends along the midline of the
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Spinal Cord posteriorly.
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The spinal cord can be divided into spinal
cord segments.
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One spinal cord segment gives rise to the
anterior and posterior nerve roots
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which comes together to form a spinal
nerve on each side of the Spinal Cord.
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There are thirty-one Spinal Cord segments:
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eight Cervical, twelve Thoracic,
five Lumbar, five Sacral,
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and one Coccygeal.
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A spinal nerve pair shares its name with
the spinal cord segment it arises from,
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so, for example, the spinal nerves arising
from the C1 spinal cord segment are named
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the C1 spinal nerves, logically the number
of paired spinal nerves is the same as the
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number of Spinal Cord segments.
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Now, not all regions of the Spinal Cord
are the same size, there are two regions
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that carry more fibers and are therefore
wider, resulting in
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two Spinal Cord enlargements.
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The cervical enlargement spans the C4
through T1 segments.
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The anterior rami of the spinal nerves
that arise from most of these segments,
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form the Brachial Plexus, which provides
innervation for the upper limbs.
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The Lumbosacral enlargement spans the T11
through S1 segments.
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The anterior rami of the spinal nerves
that arise from most of these segments,
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form the Lumbar and the Sacral Plexuses,
which provide innervation
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for the lower limbs.
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Let's take a closer look at spinal nerves.
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Spinal nerve roots travel laterally
from their spinal cord segment through
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the vertebral canal to their respective
opening which for most spinal nerves is
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an intervertebral foramen.
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Just before reaching the opening,
the roots unit to create a spinal nerve,
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which then exits the vertebral canal.
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Now, let's look at how each pair of spinal
nerves exit the vertebral collum.
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The C1 spinal nerve exits above the
arch of the C1 vertebra.
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Nerves C2 to C7, exit through the
Intervertebral Foramina superior
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to their corresponding vertebra, while
C8 exits through the
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intervertebral foramina between the C7
and the T1 vertebrate.
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Nerves T1 to L5 then exit through
intervertebral foramina inferior to
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their corresponding vertebra.
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The S1 to S4 spinal nerves branch into
anterior and posterior rami and then exit
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through the anterior and posterior
sacral foramina inferior to their
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corresponding vertebra.
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And finally, the S5 and Coccygeal nerves
exit through the Sacral hiatus, which is
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inferior to the S5 vertebra.
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Now, since the Spinal Cord only extends to
the L1, L2 vertebral level,
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the Lumbar, Sacral and Coccygeal nerve
roots travel inferiorly through the
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remaining vertebral canal to reach their
respective openings.
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In doing so, they form a bundle of nerve
roots inferior to the Spinal Cord called
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the Cauda Equina.
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Fun fact, Cauda Equina is Latin for horses
tail due to the fact that this bundle
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of spinal nerve roots resembles the hair
of a horses tail.
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[Horse Sound]
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There is also another important
structure within the distal portion of the
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vertebral canal called the Filum Terminale.
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It extends from the Conus Medullaris and
descends among the nerve roots of the
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Cauda Equina to attach to the Dorsum
of the Coccyx, acting as an anchor for the
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spinal cord within the vertebral canal.
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Okay, now let's take a short break, can
you label these images?
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Now, the Spinal Cord and the spinal nerve
roots are covered by three membranes
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collectively called the Spinal Meninges;
the spinal Dura Mater,
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Arachnoid Mater, and Pia Mater.
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The Spinal Meninges are continuous
with the Cranial Meninges through the
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Foramen Magnum and they function to
support and protect the Spinal Cord,
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they also contain the Cerebrospinal Fluid
or CSF for short
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in which the Spinal Cord is suspended.
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Now, let's talk a little bit more about
each meningeal layer.
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The Dura Mater is the outer most
layer of the spinal meninges.
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This fibrous layer forms the tubular
sheath along the vertebral canal called
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the Dural Sac.
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The space that separates the Dural Sac
from the bony walls of the vertebral canal
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is called the Epidural Space and it
contains the internal vertebral
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Venus Plexus surrounded by epidural fat.
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Superiorly, both the Dural Sac and
epidural space extend to the foramen
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magnum where the spinal Dura Mater
is continuous with the cranial Dura Mater.
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Inferiorly, the Dural Sac and Epidural
Space terminates at the
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level of the S2 vertebra.
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The Dural Sac is also anchored too the
coccyx via the Filum Terminale,
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The Dura Mater also has tapered lateral
extensions called the Dural root sheaths,
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which cover the anterior and posterior
spinal nerve roots that arise from the
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Spinal Cord segments.
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At the intervertebral foramina, these
sheaths merge with the outer covering of
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the spinal nerves called the Epineurium.
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The spinal Dura Mater is innervated
by branches of the spinal nerves called
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the recurrent meningeal nerves.
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Next is the Arachnoid Mater, which lines
the spinal Dura Mater and
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the dural root sheaths internally.
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It encloses the subarachnoid space
which is filled with CSF.
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Inferiorly, the subarachnoid space extends
beyond the Conus Medullaris
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of the spinal cord.
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The space inferior to the Conus
Medullaris which extends to the L2 vertebra
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to the S2 vertebra, is called the Lumbar
Cistern.
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It contains cerebral spinal fluid and the
Cauda Equina.
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Now, the Arachnoid Mater is not
actually attached to the Dura, but rather
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the CSF within the Subarachnoid space
presses the Arachnoid Mater
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against the Dura.
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Usually there is no space between the two
layers, however, if bleeding occurs, then
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blood can push them apart creating a
potential subdural space.
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In an anatomy lab, you might notice that
the Arachnoid Mater is separated
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from the Dura mater, this happens in
cadavers because there is not CSF in
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the Subarachnoid space is to press the
arachnoid against the Dura.
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The Arachnoid Mater is separated from
the Pia Mater by CSF, but connective
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tissue strands called Arachnoid Trabeculae
make connections between the
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Arachnoid and Pia Mater through the
Subarachnoid space.
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The Pia Mater is the inner most meningeal
layer which covers the Spinal Cord,
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spinal nerve roots and spinal blood vessels.
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A fine thread of the spinal cord Pia Mater
forms the Filum Terminale,
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extending inferiorly from the Conus
Medullaris.
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Laterally, the Pia Mater forms extensions
along the Spinal Cord
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called the Denticulate Ligaments.
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These ligaments arise between the anterior
and posterior nerve roots on both sides
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of the spinal cord.
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The Denticulate Ligaments have triangular
processes that extend laterally and pass
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through the Arachnoid to attach to the
Dura Mater.
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Superiorly, the Denticulate Ligaments
attach to the Cranial Dura Mater.
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The Denticulate Ligaments along with the
Filum Terminale help suspend the
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spinal cord in the Cerebrospinal Fluid of
the Subarachnoid Space.
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Okay, now that we've discussed the
Meninges, we can take a short break.
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Can you recall which structures anchor
Spinal Cord?
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The blood supply for the spinal cord comes
from a few sources.
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First, there are three long longitudinal
arteries, one anterior spinal artery,
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and two posterior spinal arteries.
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The anterior spinal artery originates
within the cranium from branches
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of the vertebral arteries.
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It extends inferiorly along the
interior Median Fissure.
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Its branches, called the Sulcal arteries,
enter the Spinal Cord through the
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anterior Median Fissure.
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The two posterior spinal arteries
also originate in the Cranium
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either as branches of the Vertebral
arteries or the posterior inferior
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Cerebellar arteries.
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The three longitudinal arteries can only
supply the superior parts of the
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Spinal Cord by themselves.
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The circulation to the majority of the
Spinal Cord depends on the
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posterior and anterior Segmental
Medullary arteries and Radicular arteries
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that run along the spinal nerve roots.
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The posterior and anterior Segmental
Medullary arteries arise at various
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vertebral levels from the spinal branches
of the following arteries:
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Ascending Cervical, Deep Cervical,
Vertebral, Posterior Intercostal,
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and Lumbar Arteries.
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These arteries supply blood mainly
to the Cervical and
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Lumbosacral Enlargements.
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Additionally, a branch of an Inferior
Posterior Intercostal or
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Upper Lumbar Artery, called the Great
Anterior Segmental Medullary Artery,
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also reinforces the blood supply to the
lower two-thirds of the Spinal Cord.
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Lastly, branches also give rise to the
Radicular arteries which are small
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arteries that supply blood to the
spinal nerve roots and their coverings.
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Unlike the segmental arteries, they do not
anastomose with the anterior and
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posterior spinal arteries.
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Finally, let's discuss the veins of the
Spinal Cord.
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The veins of the Spinal Cord usually
follow the arteries.
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There are typically three anterior
and three posterior spinal veins.
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They drain into anterior and posterior
Medullary and Radicular veins which
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communicate with the Epidural Venus
Plexus.
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From the Epidural Venus Plexus,
blood flows to the Dural sinuses of the
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Cranium, the Vertebral veins and the
External Venus Plexuses.
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Alright, as a quick recap, the Spinal Cord
has thirty-one spinal segments and pairs
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of spinal nerves: eight Cervical, twelve
Thoracic, five Lumbar, five Sacral, and
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one Coccygeal.
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The two enlargements of the Spinal
Cord are the Cervical Enlargement and
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Lumbosacral Enlargement.
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The Spinal Cord and the spinal nerve roots
are covered by the spinal meninges, which
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consist of the Dural Mater, Arachnoid
Mater and Pia Mater.
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The Dura Mater is separated from the bony
walls of the Vertebral Canal
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by the Epidural Space, the spinal
Arachnoid Mater is pressed against the
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Dura by the Cerebrospinal Fluid
of the Subarachnoid Space.
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The spinal Pia Mater directly covers the
Spinal Cord, the spinal nerve roots
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and spinal blood vessels.
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Inferior to the Spinal Cord, it
continues as a fine thread
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called the Filum Terminale.
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Laterally the spinal Pia Mater forms
extensions along the Spinal Cord called
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the Denticulate Ligaments.
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Finally, the Spinal Cord blood supply
comes from the anterior spinal artery,
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the two posterior spinal arteries, and the
posterior anterior segmental medullary
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arteries, including the Great Anterior
Segmental Medullary Artery.
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Venous blood drains through the three
anterior spinal veins
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and three posterior spinal veins.
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