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The original keystone refers to a wedge-shaped
slab at the peak of a stone arch.
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If you remove the keystone, you risk the whole
thing tumbling down.
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And some ecosystems work the same way, relying
on one keystone species.
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In biology, keystone species have a disproportionately
large impact on their habitat, given their
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size or numbers.
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Their daily activities affect all kinds of
other species, directly or indirectly.
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And a change in their number can cause a trophic
cascade, where effects ripple throughout the
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ecosystem, often in surprising ways.
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Take the grey wolves of Yellowstone National
Park in Wyoming.
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Their numbers dropped to zero around the early
1930s, mostly because of hunting.
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But in 1995, wolves were reintroduced to the
park, and the wolves’ impact has been monitored
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ever since.
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Turns out, that impact is seriously important.
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The wolves’ absence and reintroduction changed
Yellowstone in large and often surprising
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ways, right down to how the rivers flow.
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For one thing, wolves prey on elk — and
the elk know it!
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So their behavior changes when wolves are
in the area.
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Elk are generally nomadic, staying on the
move as they munch on plants.
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But without any wolves around, the elk became
much less cautious, preferring to settle down
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and eat a lot in one place at a time.
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The elk even ate all the way down to the riverbank
– somewhere they wouldn’t normally dare
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linger if there were wolves around.
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Instead of eating a little bit from lots of
different plants, they ate so much at a time
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that aspen and cottonwood trees declined,
and so did the riverside willows that beavers
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and some songbirds depend on.
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And beavers are also keystone species, because
of their special engineering talents.
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Their dams slow the flow of rivers, preventing
floods and offering a wide range of habitats
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for all kinds of species.
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Ever since wolves were reintroduced to Yellowstone,
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willows have grown better, beaver colonies
have increased from just one to nine, and
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songbirds are flourishing too.
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Elk numbers declined at first, but now seem
to have stabilized.
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And bison numbers are up, probably because
of a decrease in competition from the elk.
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Research is ongoing to assess the wolves’
long-term impact, considering the many subtle
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interconnections in this complex ecosystem.
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But as top predators, it’s clear that the
presence, absence, and reintroduction of wolves
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has resonated throughout Yellowstone.
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And research into other wolf habitats will
help work out what’s unique to Yellowstone
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and what can be applied elsewhere.
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Elephants also do a lot to manage their environment
— from what goes on in their mouths, to
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what comes out the other end.
-
They’re sometimes known as the “mega-gardeners
of the forest”.
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For example, without African Forest elephants,
a single species of acacia tree tends to dominate
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in African forests.
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It grows fast and shuts out the light from
other plants.
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And the elephants help counter that.
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They sometimes knock down acacia in their
search for food, opening up space and a critical
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shaft for light.
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Other plant species seize their chance, adding
to the forest’s biodiversity.
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The smaller branches they knock down provide
hidey-holes for lizards, which are more diverse
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in places where elephants roam.
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When reaching for leaves or fruit, they’ll
often knock off a bunch more.
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And these windfalls get picked up by smaller
ground-dwellers like warthogs, as well as
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kudu, which is a type of antelope.
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All this eating results in plenty of dung
– around a metric ton every week.
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And all that poop becomes a mini-ecosystem
all by itself, since it’s rich in nutrients
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the elephants couldn’t process.
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Fungi live in it, as do insects like beetle
larvae, crickets and spiders.
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In 2009, three species of frog were found
happily living in Asian elephant dung.
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And of course, it’s excellent fertilizer.
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Many plant species grow better in elephant
dung than in poop from any other animal.
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Their wide roaming and variation in movements
make them especially good at spreading seeds
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to new places.
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But you don’t have to be as big as an elephant
to have a jumbo-sized impact on your ecosystem.
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Parrotfish are named for their tough, almost
beak-like mouthparts.
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These hardened lips are well adapted for plucking
algae off coral, their main food source in
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their warm reef environment.
-
It may not be glamorous, but it turns out
to be vital to keeping the reef healthy.
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Without parrotfish, macroalgae can smother
the coral to the point of killing it off.
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And no coral means no reef, which would be
big trouble for the species that call coral
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reefs home.
-
A major report covering four decades of research
in the Caribbean identified parrotfish as
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a group to keep a special eye on.
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Coral reefs are generally having a tough time
of it: with climate change, ocean acidification
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and pollution steadily degrading the ecosystem.
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The parrotfish’s maintenance work helps
keep reefs resilient, so they can recover
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from blows like sudden heating or a hurricane.
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Caribbean reefs where parrotfish are heavily
fished are suffering
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the most.
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So, Like all keystone species, they really
hold their ecosystems together.
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